Epstein Secrets

t much larger than the degree, as well as the phenotype space. 2.5.1 The well-mixed population As first example we consider the frequency dependent Moran process in a well mixed population of size N (Nowak et al 2004, Taylor et al 2004, Nowak 2006a) (Fig.la). In the language of evolutionary graph the

ho-interacts-with-whom changes during the evolutionary process (Fig 2.2). For mathematical convenience we con- sider evolutionary game dynamics in a Wright-Fisher process with constant population size [63]. A frequency dependent Moran process [110] or a pairwise comparison process [156], which is more realist

t in common, they interact several times. An interaction is given by the usual payoff matrix. The system evolves according to synchronous updating (Wright-Fisher process). There are discrete, non-overlapping generations. All individuals update at the same time. The population size is constant. Individuals re

weak selection (as opposed to 6 cc 1) shall become clear at the end of this section. The dynamics of the system is given by the frequency dependent Moran process. In each time step a randomly chosen individual is replaced by a copy of an individual chosen with probability proportional to its payoff.

Evolutionary updating occurs according to the frequency dependent Moran process (Nowak et al, 2004; Taylor et al, 2004) or the frequency dependent Wright-Fisher process (Imhof & Nowak, 2006). Reproduction is proportional to fitness and subject to a mutation rate u. With probability 1 — u the offspring inheri

A., 2005. Evolutionary cycles of coop- eration and defection. PNAS, 102, 1079710800. [63] Imhof LA, Nowak MA, 2006. Evolutionary game dynamics in a Wright-Fisher process. J. Math. Biol 52, 667-681. [64] Jansen VA, van Baalen M, 2006. Altruism through beard chromodynkunics. Nature, 440, 663-666. [65] Kandor

ure 438: T148-T150. 64. Claussen J, Traulsen A (2005). Nongaussian fluctuations arising from finite populations: Exact results for the evolutionary Moran process. Phys Rev E 71: 025101. 65. Docbeli M, C Hauert (2005). Models of cooperation based on the Prisoner's Dilemma and the Snowdrift game. Ecol

nergy and discounting of cooperation in social dilemmas. I theor Biol 239: 195-202. 125. Imhof LA, MA Nowak (2006). Evolutionary game dynamics in a Wright-Fisher process. Math Biol 52: 667-681. 126. Iwasa Y, MA Nowak, F Michor (2006). Evolution of resistance during clonal expansion. Genetics 172: 2557-2566.