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1992, 2002, Hofbauer 1999). Evolutionary graph theory is an extension of spatial games to more general popu- lation structures and social networks (Lieberman et al 2005, Ohtsuki et al 2006a,b, Pacheco et al 2006, Szabo & Fath 2007, Taylor et al 2007a, Santos at al 2008, RI et al 2008). The members of the
ho-interacts-with-whom changes during the evolutionary process (Fig 2.2). For mathematical convenience we con- sider evolutionary game dynamics in a Wright-Fisher process with constant population size [63]. A frequency dependent Moran process [110] or a pairwise comparison process [156], which is more realist
t in common, they interact several times. An interaction is given by the usual payoff matrix. The system evolves according to synchronous updating (Wright-Fisher process). There are discrete, non-overlapping generations. All individuals update at the same time. The population size is constant. Individuals re
Science, 303, 793-799. 0.14] Nowak, M.A., Sigmund, K., 2005. Evolution of indirect reciprocity. Nature 427, 1291- 1298. (115] Ohtsuki H, Hauert C, Lieberman E, Nowak MA, 2006a. A simple nile for the evolution of cooperation on graphs and social networks. Nature, 441, 502-505. [116] Ohtsuki, H., Iwasa, Y
Evolutionary updating occurs according to the frequency dependent Moran process (Nowak et al, 2004; Taylor et al, 2004) or the frequency dependent Wright-Fisher process (Imhof & Nowak, 2006). Reproduction is proportional to fitness and subject to a mutation rate u. With probability 1 — u the offspring inheri
A., 2005. Evolutionary cycles of coop- eration and defection. PNAS, 102, 1079710800. [63] Imhof LA, Nowak MA, 2006. Evolutionary game dynamics in a Wright-Fisher process. J. Math. Biol 52, 667-681. [64] Jansen VA, van Baalen M, 2006. Altruism through beard chromodynkunics. Nature, 440, 663-666. [65] Kandor
r, C Taylor (2005). Grammar structure and the dynamics of language evolution. In Proceedings of the 8th European Conference on Artificial Lifc. 80. Lieberman E, C Hauert, MA Nowak (2005). Evolutionary dynamics on graphs. Nature 433: 312-316. 81. Lieberman E, C Hauert, MA Nowak (2005). Evolutionary dynami
nergy and discounting of cooperation in social dilemmas. I theor Biol 239: 195-202. 125. Imhof LA, MA Nowak (2006). Evolutionary game dynamics in a Wright-Fisher process. Math Biol 52: 667-681. 126. Iwasa Y, MA Nowak, F Michor (2006). Evolution of resistance during clonal expansion. Genetics 172: 2557-2566.
Entities connected to both Lieberman and Wright-Fisher

Martin Nowak
PERSON
George W. Bush
PERSONCourtney Wild
PERSONEvolutionary Dynamics
ORGANIZATION
Harvard University
ORGANIZATIONEmmy Taylor
PERSONTraulsen
PERSON
Morgan Stanley
ORGANIZATIONRobert Trivers
PERSONImhof
PERSON
Stanford University
LOCATIONSantos FC
LOCATION
Ellison
PERSONNowak MA
PERSONKomarova
PERSONDurrett
PERSONP. Natl
ORGANIZATIONRoyal Soc
ORGANIZATIONKerr et al 2002
ORGANIZATIONHofbauer Sc
ORGANIZATION