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(2009) Indirect reciprocity and strategic reputation building in an experimental helping game. Games (con &hate 67:399-407. 5. Mcquet ), Revert C, Traulsen A, Milins1/41 M (2011) Shame and honour drive cooper- ation. Biel Lett 7:899-901. Hilbe et el. 9NAS Latest Articles I 5 of 6 EFTA00803982 6. Oht
idge, UK). 47. Szolnoki A. Szabo G (2004) Phase transitions for rock-scissorypaper game on different networks. Mys Rev E 70037102 48. Claussen 1C, Traulsen A (2008) Cyclic dominance and biodiversity it well-mixed populations, Phys ROV Lett 100:058104. 09. Szolnoki A, et al. (2014) Cyclic dominance in e
Dynamics, Harvard University, Cambridge, MA 02138; 'Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138; and °Department of Mathematics, Harvard University, Cambridge, MA 02138 Edited by Brian Skyrms, University of California, Irvine, CA, and approved October 16, 2018 (received for
Jennifer; University of British Columbia, Department of Mathematics Hauert, Christoph; University of British Columbia, Department of Mathematics Traulsen, Arne; Max-Planck-Institute for Evolutionary Biology, Research Group for Evolutionary Theory Milinski, Manfred; Max-Planck-Institute for Evolutiona
of the commons. 52 53 54 55 Keywords: cooperation, honour, shame, public goods game, tragedy of the commons 56 57 58 59 60 Jacques, Haunt, Traulsen, & Milinski http://mc.ma nuscri ptcentral.com/b I EFTA_R1_02030116 EFTA02690245 Submitted to Biology Letters Page 2 of 9 1 Jacques, Haunt, Tr
logy Letters 1 2 3 Fear of shame drives cooperation 4 5 6 Jennifer Jacquetl 2., Christoph Hauert', Arne Traulsen3 and Manfred Milinski4 7 8 Department of Mathematics, University of British Columbia, 1984 Mathematics Road, 9 10 Vancouver, British Columbia, V6T 1Z2 Canada (where the research was carried out) 11
ary dynamics (Antal et al.. 2009; Allen and Nowak. 2014; Fu et al.. 2014; Haven and Doebelt, 2004; Szabo and Kith. 2007; Antal and Scheming. 2006; Traulsen et al., 2008; van Veelen et al.. 2014; Simon ct al.. 2013) demonstrate that inclusive fitness is not needed for understanding any phenomenon in evo
l'"', Carl yeller • Raghavendra Gadagkar -1. Martin A. Nowak a•d•g.s Program for twoltalonary Dynamks. Hanard University, Cambridge, MA 02(38. USA °Department of Mathematics. Emmanuel College, Bosron. MA 02115, USA `Center for Mathematical Sciences and Applicanons. Harvard Unhersky. Cambridge, MA 02138. USA Depaytmcnr o
Entities connected to both Traulsen and Department of Mathematics

Martin Nowak
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Harvard University
ORGANIZATIONCourtney Wild
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George W. Bush
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Princeton University
ORGANIZATIONNowak MA
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R. Soc
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