Epstein Secrets

policing Jason W. Olejarz a, Benjamin Allen l'"', Carl yeller • Raghavendra Gadagkar -1. Martin A. Nowak a•d•g.s Program for twoltalonary Dynamks. Hanard University, Cambridge, MA 02(38. USA °Department of Mathematics. Emmanuel College, Bosron. MA 02115, USA `Center for Mathematical Sciences and Applicanons. Ha

tion. Fxol. Lett 19 (Ile 74-84. Gadagkar. It. 2001. 'Ilk Social Biology of Ropalidia hlargiluu: low.u4 Under- standing the Evolution of EuSOCiality. Harvard University Press. Cambridge. MA. Cadagkar. R, 2004. Why do honey bee workers destroy each other's eggs? J. Mout 29(3). 213-217. Cardneu. A.. West, SA. Wild. G.. 20

l'"', Carl yeller • Raghavendra Gadagkar -1. Martin A. Nowak a•d•g.s Program for twoltalonary Dynamks. Hanard University, Cambridge, MA 02(38. USA °Department of Mathematics. Emmanuel College, Bosron. MA 02115, USA `Center for Mathematical Sciences and Applicanons. Harvard Unhersky. Cambridge, MA 02138. USA Depaytmcnr o

y, Cambridge, MA 02138; 'Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138; and °Department of Mathematics, Harvard University, Cambridge, MA 02138 Edited by Brian Skyrms, University of California, Irvine, CA, and approved October 16, 2018 (received for review June 19, 2018)

Dynamics, Harvard University, Cambridge, MA 02138; 'Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138; and °Department of Mathematics, Harvard University, Cambridge, MA 02138 Edited by Brian Skyrms, University of California, Irvine, CA, and approved October 16, 2018 (received for

T Austria, KlostemeubLeg, Austria, 2 Program for Evolutionary Dynamics, Department of Organismic and Evolutionary Biology, Department of Mathematics, Harvard University, Cambridge, Massachusetts, United States of America Abstract A fundamental question in biology is the following: what is the time scale that is nee

lam2, Martin A. Nowak2 1 IST Austria, KlostemeubLeg, Austria, 2 Program for Evolutionary Dynamics, Department of Organismic and Evolutionary Biology, Department of Mathematics, Harvard University, Cambridge, Massachusetts, United States of America Abstract A fundamental question in biology is the following: what is the t

414. (doi: 10.1098/rsb1.2006.0509) 47 48 13 Dunbar, R. 1997 Grooming. Gossip and the Evolution of Language, Cambridge, 49 50 51 Massachusetts, Harvard University Press. 52 53 54 55 56 57 58 59 60 7 http://mc.ma nuscri ptcentral.com/b I EFTA_R1_02030122 EFTA02690251 Submitted to Biology Letters Page 8

logy Letters 1 2 3 Fear of shame drives cooperation 4 5 6 Jennifer Jacquetl 2., Christoph Hauert', Arne Traulsen3 and Manfred Milinski4 7 8 Department of Mathematics, University of British Columbia, 1984 Mathematics Road, 9 10 Vancouver, British Columbia, V6T 1Z2 Canada (where the research was carried out) 11