haffer 1988, Kandori et al 1993, Kandori & Rob 1995, Schreiber 2001, Nowak et al 2004, Taylor et al 2004, Wild and Taylor 2004, Traulsen et al 2005, Antal & Scheuring 2006, Imhof & Nowak 2006, Lessard & Ladret 2007). Evolutionary updating occurs according to the frequency dependent Moran process (Nowak et al, 2004; Ta