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plaining evolution of worker policing (Foster and Ratnieks 2001a,c: Hartmann et al_ 2003; Hammond and Keller. 2004; Wenseleers and Ratnieks. 2006b; Helantera and Sundstrom. 2007; Khila and Abouheif. 2008; Zanette et al. 2012). Yet reliable pub- lished data on the effect that policing has on colony reprodu