ak, MA, 2015. The evolution of non reproductive workers in insect colonies with haplocliplold genetics elite 4. e08918. Peters. JAL. Quelkr. D.C.. Imperatriz-Fonseca. V.L. Roulade. D.W_ Strassmann. 1999. Mate number, kin selection and social conflicts in stingless bees and honeybees, Proc. R. Soc. loud. B 266. 3