licing mutant allele. we must consider all colony types that have 0,1, or 2 mutant A alleles: these are Oa. n; aa.n —1: Ao.n: aa. n —2; Ao.n-1; and M. n. The colonies aa,n: aa.n —1: Aa.n: aa.n —2: and Mae do not contain non-police workers: the efficiency of those colonies is r,, and the fraction of