should be absent if queens are singly mated. EFTA_R1_02076880 EFTA02702876 104 1W. OhIan et al /journal of Theoretkal Bletrov 399 (2016)103-116 Woyciechowski and Lomnicki (1987) perform a calculation based on population genetics and conclude that workers should raise their nephews (sons of other workers)