olicing (Ratnieks. 1988; Ratnieks et al.. 2006; Gadagkar, 2001; Wenseleers and Ratnieks. 2006a) and worker sterility (Wilson. 1971; Hamilton. 1972; Olejarz et al., 2015) are two distinct phe- nomena that are widespread in the eusocial Hymenoptera. In addi- tion to worker policing, a subset of workers in a colon