e fitness theory and eusociality. Nature 471. E1-F9. Alkay. E. Van Cleve. J.. 2016. There is no fitness but fitness. and the lineage is its bearer. Philos Trans. R. Soc. 13371. 20150085. Allen. Et, Nowak. MA.. Wilson, E,0.. 2013. Limitations of inclusive fitness. Proc. Nail, Mad. 50. U.SA 110.20135-20139.