ical Biology 399(2016) 103-116 1O4 aRt` SN ELSEVIER Contents lists available at ScienceDirect Journal of Theoretical Biology journal homepage: Conshlark Evolution of worker policing Jason W. Olejarz a, Benjamin Allen l'"', Carl yeller • Raghavendra Gadagkar -1. Martin A. Nowak a•d•g.s Program for t