xtant basal groups of both nonvascular and vascular plants. as well as fossil plants (Strullu-Derrien et 2014), indicate that associations between Mucoromycotina and land plants are extremely ancient. During this investigation, we examined sporophytes only and it would be desirable now to study the cryptic

mples analysed. Diverse Mucoromycotina fungi colonized lycopods, sometimes occurring within the same species, and even the same plant, and six new Mucoromycotina clades were discovered (Fig. S2). Colonization rates in ferns were higher (33 out of 58 samples) and showed specificity to Glomeromycota (Fig. SI).

rhiza-like relation- ship, whereby there is reciprocal exchange of plant carbon (C) for fungal nitrogen (N) and phosphorus (P), with members of the Mucoromycotina (Bidartondo et al, 2011; Field et 2014), a fungal lineage considered basal or sister to the Glomeromycota (James etal., 2006; Lin nal, 2014). Seco