Urban Environmental Sciences, Tokyo Metropolitan University, Minami-ohsawa 1-1, Hachiohji, Tokyo 192-0397 Japan 5 EFTA_R1_01696124 EFTA02543935 Japan Society for the Promotion of Science (DC1), Ichibancho, Chiyoda-ku, Tokyo 102.8471, Japan PRESTO (Precursory Research for Embryonic Science and Technology), Japan Science and Technology