Columbia, 1984 Mathematics Road, 9 10 Vancouver, British Columbia, V6T 1Z2 Canada (where the research was carried out) 11 12 13 2Sea Around Us Project, Aquatic Ecosystem Research Laboratory, 2202 Main Mall, Vancouver, 14 15 British Columbia, V6T 1Z4 Canada 16 17 18 3Research Group for Evolutionary Theory, Max-Planck-Institute for