EFTA_R1_02030115 EFTA02690244 Page 1 of 9 Submitted to Biology Letters 1 2 3 Fear of shame drives cooperation 4 5 6 Jennifer Jacquetl 2., Christoph Hauert', Arne Traulsen3 and Manfred Milinski4 7 8 Department of Mathematics, University of British Columbia, 1984 Mathematics Road, 9 10 Vancouver, Brit