OPEN 8 ACCESS Freely available online • • • (t) -PLOS COMPUTATIONAL BIOLOGY The Time Scale of Evolutionary Innovation Krishnendu Chatterjee", Andreas Paviogiannisl , Ben Adlam2, Martin A. Nowak2 1 IST Austria, Klos

Christine Maxwell 3/24/03 Written Permission of the Author is Required for Copying or Distribution EFTA00578135 Page 23 of 25 of LINGUISTIC AND COMPUTATIONAL CONSULTANTS Bert Vaux, Philology Consultant, has been an Assistant/Associate Professor in the Linguistics Department of Harvard University since Ju