A. W. The speed of evolution and maintenance of variation in asexual populations. Carr. Biol. 17, 385-394 (2007). 4. McCandlish. D. M.. Epstein. C. L & Plotkin. I. B. Formal properties of the probability of fixation: identities, inequalities and approximations. Them. Poput Blot 99, 98-113 (2015). 5. Nowa