Epstein Secrets

tedness—including the effects of policing on a colony's rate of production of offspring—may play a role in explaining evolution of worker policing (Foster and Ratnieks 2001a,c: Hartmann et al_ 2003; Hammond and Keller. 2004; Wenseleers and Ratnieks. 2006b; Helantera and Sundstrom. 2007; Khila and Abo

Biology journal homepage: Conshlark Evolution of worker policing Jason W. Olejarz a, Benjamin Allen l'"', Carl yeller • Raghavendra Gadagkar -1. Martin A. Nowak a•d•g.s Program for twoltalonary Dynamks. Hanard University, Cambridge, MA 02(38. USA °Department of Mathematics. Emmanuel College, Bosron. MA 02115

that affect the intensity of policing, Our paper is based on an analysis of evolutionary dynamics and population genetics of haplodiploid species (Nowak et al., 2010; Olejarz et al., 2015). It does not use inclusive fitness theory. Spe- cifically, we adapt the mathematical approach that was developed

„ Jonathan D. Eric Lib* M•ONY ca tch. Express Foster, Taylor Gell-man Murray Gilman, Kenneth , EFT

Mlr Mitchell, David Mitchell, Senator George Morrison, Larry Myhrvold, Nathan INNlucfwl ve t N.A. Pro erty, Inc iddtiL0elcACIr EFTA00183021 Nowak, Martin New Albany Country Newman, Larry Esq. Ciatman, Bob Ovitz, Michael All ( Man TWIT G,00p Packer, Mark Pagano, Joe idn P man ltzhak P

Research/University of Cambridge, 119-129. 116. Foster KR (2006). The Phoenix effect. Nature 441: 291-29

s: Cooperation and bifurcation. Theor Popul Biol 73: 257-263. 208. Jones S, W Chen, G Parmigiani, F Diehl, N Beerenwinkel, T Antal, A Traulsen, MA Nowak, C Siegel, VE Velculescu, KW Kinzler, B Vogelstein, J Willis, SD Markowitz (2008). Comparative lesion sequencing provides insights into tumor evolut